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Before 2.6 million years ago (Ma), no archaeological record has been securely documented, and therefore there is no evidence of hominin tool use. Then, at 2.6–2.58 Ma, there is widespread evidence for tool manufacture and use at several archaeological sites, with undisputed stone tools and fossil fauna at Gona, Ethiopia. Additionally, the evidence from the earliest archaeological sites at Gona shows that the earliest stone tool makers were skilled flintknappers and were able to select high quality stone raw materials. The possible reasons behind this seeming abrupt transition from the absence of stone tools to the presence thereof include sampling biases, paleogeographic influences, gaps in the geological record, paleoenvironmental change, and changes in the record of hominin evolution.Based on our observations at Gona, the earliest use of flaked stone tools is likely to be slightly older than 2.6 Ma. These stone tools represent a significant change in behavior that set the stage for subsequent hominin evolution. The paleogeographic and paleoenvironmental evidence points to the earliest use of stone tools in certain settings: usually (but not always) close to raw material sources, and ecotones between riparian woodlands and open grasslands. The earliest stone tool makers were proficient, selective, and flexible in their reduction strategies. The variability we see in Pliocene artifact assemblages has much to do with different raw material sources.

The phrase “Developed Oldowan” (DO) was originally coined by M. Leakey to describe a technologically “advanced Oldowan” artifact tradition, that preceded the Acheulian Industry. M. Leakey further identified three stages of the DO which she labeled as the DOA, DOB and DOC. The DO (sensu lato) has been generally recognized as transitional to the Acheulian, but the status of the DOB and the DOC remains unclear. In addition to a lack of clarity in terms of classification, the DO also suffers from a lack of secure radiometric dates, even at Olduvai where it was first identified. Despite such shortcomings, archaeologists still assign assemblages into the DO, as supposedly “intermediate” or transitional between the Oldowan and the Acheulian. However, a closer look at the DO assemblages from Olduvai Gorge and other sites in Africa and the Middle East shows that the artifacts assigned into this tradition are not technologically drastically different from the preceding Oldowan. Probably the flaking characteristics of the raw material types (e.g., quartzite and limestone, and to a lesser extent basalt) and the original shape of the cobbles used by hominins may have played a major role in the final shape of the “distinctive” artifact types (such as spheroids/subspheroids) used for assigning assemblages into the DO. Further, both the DOB and the Acheulian appeared ˜1.7 million years ago (Ma) in the archaeological record, making it unlikely that the DO is a transitional artifact tradition that preceded the Acheulian. Our preliminary evaluation of the archaeological record at Gona, Ethiopia and elsewhere suggests a fairly abrupt appearance of the Acheulian after a temporally rapid transition from the Oldowan.

Newly recorded archaeological sites at Gona (Afar, Ethiopia) preserve both stone tools and faunal remains. These sites have also yielded the largest sample of cutmarked bones known from the time interval 2.58-2.1 million years ago (Ma). Most of the cutmarks on the Gona fauna possess obvious macroscopic (e.g., deep V-shaped cross-sections) and microscopic (e.g., internal microstriations, Herzian cones, shoulder effects) features that allow us to identify them confidently as instances of stone tool-imparted damage caused by hominid butchery. In addition, preliminary observations of the anatomical placement of cutmarks on several of the recovered bone specimens suggest that Gona hominids may have eviscerated carcasses and defleshed the fully muscled upper and intermediate limb bones of ungulates-activities that further suggest that Late Pliocene hominids may have gained early access to large mammal carcasses. These observations support the hypothesis that the earliest stone artifacts functioned primarily as butchery tools and also imply that hunting and/or aggressive scavenging of large ungulate carcasses may have been part of the behavioral repertoire of hominids by c. 2.5 Ma, although a larger sample of cutmarked bone specimens is necessary to support the latter inference. (c) 2004 Elsevier Ltd. All rights reserved.

Published evidence of Oldowan stone exploitation generally supports the conclusion that patterns of raw material use mere determined by local availability. This is contradicted by the results of systematic studies of raw material availability and use among the earliest known archaeological sites from Gona, Afar, Ethiopia. Artifact assemblages from six Pliocene archaeological sites were compared with six random cobble samples taken from associated conglomerates that record pene-contemporaneous raw material availability. Artifacts and cobbles were evaluated according to four variables intended to capture major elements of material quality: rock type, phenocryst percentage, average phenocryst size, and groundmass texture. Analyses of these variables provide evidence of hominid selectivity for raw material quality. These results demonstrate that raw material selectivity was a potential component of Oldowan technological organization from its earliest appearance and document a level of technological sophistication that is not always attributed to Pliocene hominids. (c) 2004 Elsevier Ltd. All rights reserved.

A 4.5 Ma record of fluvial and lacustrine deposits is well exposed at Gona, in the Afar Depression of Ethiopia. We use isotopic values of pedogenic carbonate and fossil teeth to reconstruct Plio-Pleistocene environmental change at Gona. An increase in δ13C values of pedogenic carbonates since 4.5 Ma points to a shift from woodlands to grassy woodlands in the early Pliocene, -10.4 to -3.9‰ (VPDB), to more open but still mixed environments in the late Pleistocene, -3.0 to -1.4‰ (VPDB). This pattern is also seen in isotopic records elsewhere in East Africa. However, at 1.5 Ma the higher proportion of C4 grasses at Gona is largely a result of a local facies shift to more water-limited environments. The wide range of δ13C values of pedogenic carbonate within single stratigraphic levels indicates a mosaic of vegetation for all time intervals at Gona that depends on depositional environment. Elements of this mosaic are reflected in δ13C values of both modern plants and soil organic matter and Plio-Pleistocene soil carbonate, indicating higher amounts of C4 grasses with greater distance from a river channel in both the modern and ancient Awash River systems. δ18O values of pedogenic carbonates increase up-section from -11.9‰ in the early Pliocene to -6.4‰ (VPDB) in the late Pleistocene. The wide range of δ18O values in paleovertisol carbonates from all stratigraphic levels probably reflects short-term climate changes and periods of strong evaporation throughout the record. Based on the comparison between δ18O values of Plio-Pleistocene pedogenic carbonates and modern waters, we estimate that there has been a 6.5‰ increase in mean annual δ18O values of meteoric water since 4.5 Ma. δ18O values of pedogenic carbonate from other East African records indicate a similar shift. Increasing aridity and fluctuations in the timing and source of rainfall are likely responsible for the changes in δ18O values of East African pedogenic carbonates through the Plio-Pleistocene. © 2004 Elsevier B.V. All rights reserved.

Fluvio-lacustrine sediments of the Hadar and Busidima Formations along the northern Awash River (Ethiopia) archive almost three million years (3.4 to <0.6 Ma) of human evolution, including the earliest documented record of stone toolmaking at 2.5-2.6 Ma. This paper brings together sedimentologic and isotopic evidence for the paleoenvironmental context of early hominids from both formations, but with particular emphasis on the setting for the early toolmakers. The pre-2.92 Ma record (Hadar Formation) is characterized by low-gradient fluvial, paludal, and lacustrine deposition in an undissected topography most analogous to reaches of the modern middle Awash River near Gewane. The Gona area experienced repeated deep dissection and aggradation by the Awash River, starting between 2.92 and ca. 2.7 Ma and continuing through the top of the record at <0.6 Ma (Busidima Formation). Each aggradational succession is 10-20 m in thickness and fines upward from well-rounded conglomerates at the base to capping paleosols at the top. During this period the ancestral Awash represented by these fining upward sequences was dominantly meandering and flowed northeast, as it does today. Smaller channels tributary to the axial Awash system are also extensively exposed in the Busidima Formation. Compared to the axial-system conglomerates, the tributary channels transported finer, less mature volcanic clasts mixed with abundant carbonate nodules reworked from adjacent badlands. Stone artifacts (Oldowan; 2.6-2.0 Ma) at the oldest archaeological sites are only associated with the axial Awash system, in the bedded silts or capping paleosols of the fining upward sequences. The implements were made from rounded cobbles from the channels, but manufacture and use of the tools was always away from the channel bars, on the nearby sandy banks and silt-dominated floodplains. Archaeological sites higher in the record (Acheulian; <1.7 Ma) occur in similar axial river contexts, as well as along tributary channels further removed from artifact raw material sources. Mature paleosols in the Hadar and Busidima Formations are mostly pale to darkbrown Vertisols typified by abundant clay slickensides, pseudo-anticlinal and vertical fracturing, and carbonate nodules. Such calcic Vertisols are common in the region today, demonstrating that the paleoclimate over the past 3.4 m.y. has been semi-arid and strongly seasonal. Carbon isotopic results from pedogenic carbonates in the Vertisols allow reconstruction of the proportion of C3 plants (trees and shrubs) to C4 plants (grasses) through time. The δ 13C results from the Hadar Formation prior to 2.9 Ma range from -9.3‰ to -4.1‰, indicating a dominantly forested environment but with locally substantial (average 34%) grasses on the Awash floodplain. The δ13C values from soil carbonate in the lower Busidima Formation (2.7-1.6 Ma) increase (-6.5‰ to -2.7‰) in floodplain paleosols, indicating ∼ 50% average grass cover. Vertisols of the upper Busidima Formation (< 1.6 Ma) formed on gently sloping alluvial fans adjacent to the Awash floodplain and display even more positive δ13C values, up to -1.8‰, showing that grassland dominated the margins of the active Awash floodplain. © 2004 Geological Society of America.

Comparative biomolecular studies suggest that the last common ancestor of humans and chimpanzees, our closest living relatives, lived during the Late Miocene-Early Pliocene 1,2. Fossil evidence of Late Miocene-Early Pliocene hominid evolution is rare and limited to a few sites in Ethiopia 3-5, Kenya 6 and Chad 7. Here we report new Early Pliocene hominid discoveries and their palaeoenvironmental context from the fossiliferous deposits of As Duma, Gona Western Margin (GWM), Afar, Ethiopia. The hominid dental anatomy (occlusal enamel thickness, absolute and relative size of the first and second lower molar crowns, and premolar crown and radicular anatomy) indicates attribution to Ardipithecus ramidus. The combined radioisotopic and palaeomagnetic data suggest an age of between 4.51 and 4.32 million years for the hominid finds at As Duma. Diverse sources of data (sedimentology, faunal composition, ecomorphological variables and stable carbon isotopic evidence from the palaeosols and fossil tooth enamel) indicate that the Early Pliocene As Duma sediments sample a moderate rainfall woodland and woodland/grassland., (C) 2005 Nature Publishing Group