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Lyonsia hyalina Conrad possesses “radial mantle glands” of a complex structure located deep within the periostracal groove. They occur along the mantle edge in alignment with slightly raised striations of the periostracum, are deeply staining and are composed of three cell types. Secretory and supportive cells, which are flask-shaped, alternate throughout the gland, while a third cell type is ovoid and borders the gland. A sulfated mucopolysaccharide is secreted over the periostracum by the glands and functions in adhesion of sand grains to the shell.
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Spiral and near-spiral growth patterns of the peripheral crossed-lamellar shell microstructure of the bivalve Corbicula cf. fluminea have been discovered. This is the first report of spiral microstructures outside of nacreous and "foliated calcite" deposits. The spiral growth surface in this corbiculid is underlain by a conical crossed-lamellar microstructure. This form of lath deposition is usually associated with high concentrations of organic material located beneath the undertucked periostracum along the valve's periphery. These structures can offer the bivalve additional resistance against predators that enter the valves through the shell's edge. Similar spiral structures have not been found in all "white" forms of North American Corbicula, in the "purple" form of North American Corbicula, nor in the corbiculids Polymesoda caroliniana and P. (Geloina) erosa.
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