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A precision measurement by AMS of the antiproton flux and the antiproton-to-proton flux ratio in primary cosmic rays in the absolute rigidity range from 1 to 450 GV is presented based on 3.49×1⁢05 antiproton events and 2.42×1⁢09 proton events. The fluxes and flux ratios of charged elementary particles in cosmic rays are also presented. In the absolute rigidity range ∼60 to ∼500 GV, the antiproton ¯𝑝, proton 𝑝, and positron 𝑒+ fluxes are found to have nearly identical rigidity dependence and the electron 𝑒− flux exhibits a different rigidity dependence. Below 60 GV, the (¯𝑝/𝑝), (¯𝑝/𝑒+), and (𝑝/𝑒+) flux ratios each reaches a maximum. From ∼60 to ∼500 GV, the (¯𝑝/𝑝), (¯𝑝/𝑒+), and (𝑝/𝑒+) flux ratios show no rigidity dependence. These are new observations of the properties of elementary particles in the cosmos.